On the diversity and richness of understory bryophytes at Nectandra Cloud Forest Reserve, Costa Rica

Research output: Contribution to journalArticle


Background A survey of the understory bryophytes in the Nectandra Cloud Forest Preserve yielded 1083 specimens distributed among 55 families, represented by 74 genera of mosses, 75 genera of liverworts and 3 of hornworts. We studied and analyzed the bryophytic distribution on six types of substrates: 1) corticolous, 2) epiphyllous, 3) saxicolous, 4) terricolous, 5) aquatic and 6) lignicolous. The richness and composition of bryophyte genera are compared to those of other previous bryophyte surveys from 4 other sites with different oceanic exposures, climatic and geographic conditions in Costa Rica. New information This is a report of the first extensive general survey of bryophytes at the Nectandra Reserve, a premontane cloud forest located on the Atlantic slope of Costa Rica, an area much less studied compared to the Monteverde cloud forest on the Pacific slope.


  • Daniel H. Norris
  • Ekaphan Kraichak
  • Allen C. Risk
  • Diane Lucas
  • Dorothy J. Allard
  • Frida Rosengren
  • Theresa A. Clark
  • Nicole Fenton
  • Michael Tessler
  • Nonkululo Phephu
  • Evelyne T. Lennette
External organisations
  • University of California, Berkeley
  • Morehead State University
  • University of Vermont
  • University of Nevada, Las Vegas
  • American Museum of Natural History
  • Kasetsart University
  • Kent State University
  • Université du Québec en Abitibi-Témiscamingue
  • Tshwane University of Technology
  • Nectandra Institute
Research areas and keywords

Subject classification (UKÄ)

  • Botany


  • Bryophyte surveys, Costa Rica, Liverworts, Mosses, Nectandra Cloud Forest Reserve, Premontane cloud forest, Tropical bryophytes
Original languageEnglish
Article numbere11778
JournalBiodiversity Data Journal
Issue number1
Publication statusPublished - 2017
Publication categoryResearch