The first record of Hainosaurus (Reptilia : Mosasauridae) from Sweden
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Isolated marginal tooth crowns of the early Campanian mosasaur Hainosaurus Dollo, 1885, from the Kristianstad Basin and the Vomb Trough. southern Sweden, are described and illustrated. The teeth have been collected from a narrow stratigraphic interval corresponding to the highest belemnite zone in the lower part of the European two-fold division of the Campanian stage. A reexamination of dental and skeletal characters in two alleged species of Hainosaurus, 'H.' pembinensis Nicholls, 1988 and 'H.' gaudryi (Thevenin, 1896). and detailed comparisons with the corresponding elements in H. bernardi Dollo, 1885 and Tylosaurus proriger (Cope. 1869a), strongly indicate that 'H.' pembinensis and 'H.' gaudryi are both Tylosaurus Marsh, 1872. Diagnostic features of Hainosaurus include a very small infrastapedial process on the quadrate (conspicuous protuberance in Tylosaurus), flattened, symmetrically bicarinate marginal teeth (asymmetric and conical in Tylosaurus), short and wide pygal centra, and anteriorly situated intermediate caudal vertebral centra with dorsoventrally thin transverse processes (markedly triangular centra and thick processes in Tylosaurus).